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DNA methyltransferase 1-mediated modifications exhibit bidirectional effects: hypermethylation of promoter regions in protective genes, such as ACTN4 (encoding -actinin-4), suppresses their expression, while hypomethylation of pro-pathogenic genes, including matrix metalloproteinase-9, enhances their transcription (67, 68)

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D P Jones, R J Coates, E W Flagg, J W Eley, G Block, R S Greenberg, E W Gunter, B Jackson - Glutathione in foods listed in the National Cancer Institute's Health Habits and History Food Frequency Questionnaire - Nutr Cancer - 1992